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Application summary Application title This should be the title of your proposed project. Proposed start date Name of administering organisation Please enter the name of the organisation where you intend to hold the award. If your application is successful, this is the organisation that will be responsible for administering the award. Lead applicant's address at administering organisation Department/Division Organisation Street City/Town Postcode/Zipcode Please enter the address where you will be working at the administering organisation. If your application is successful, this is the address that will be used in the award letter. Research funding area Please select from the drop-down list the funding area that you consider your research falls under The research funding area selected is used to automatically route your application form to the appropriate Wellcome Trust grants team when it arrives at the Trust. Please note that, when received, we may reallocate your application to another research funding area if we consider it appropriate. Lead applicant Lead applicant details Full Name Department Division Page 1 of 5

Organisation Address Line 1 City/Town Postcode Telephone No. Email Address Education/training From To Qualification Subject Organisation Please provide details of relevant education/training, listing the most recent first. Career history (current/most recent first) From To Position Organisation Please provide details of your current position (if applicable) and all previous posts held, listing most recent first. Career breaks Have you had any career breaks or periods of part-time work, for example parental or long-term sick leave? We encourage applications from researchers who have taken career breaks, and wish to ensure that any such breaks are duly taken into account when considering your track record. Please state when and for what period of time you took a break, or were working on a part-time basis. Please provide details Do you wish to undertake this award part time? If you wish to undertake this award part time, you must be employed on a part-time basis. Please contact the Trust to discuss your requirements. Career contributions What are your most important research-related contributions to date? This may include contributions to health policy or practice, or to technology or product discovery and development. (350 words max.) Page 2 of 5

The examples you choose can be taken from any stage of your research career. In each case, please state what the achievement was, when it came about, why you think it is important and what impact it has had. Personal statement Describe how this Fellowship would further your research and career aspirations, and its context in your longer-term vision. How would the Fellowship differ from a more traditional postdoctoral position and how do you propose to take advantage of the opportunities offered by the scheme? (300 words max.) Research outputs List up to 20 of your most significant research outputs, ensurimg that at least five of these are from the last five years. For 10 of these outputs, provide a statement describing their significance and your contribution (up to 50 words per output). Research outputs may include (but are not limited to): Peer-reviewed publications and preprints Datasets, software and research materials Inventions, patents and commercial activity Please give citation in full, including title of paper and all authors* Citations to preprints should state Preprint, the repository name and the articles persistent identifier (e.g DOI). (*All authors, unless more than 10, in which case please use et al, ensuring that your position as author remains clear.) You should include here systematic reviews (e.g. Cochrane Reviews) and meta analyses, but exclude abstracts and literature reviews. We encourage the inclusion of articles published via open research publishing platforms, such as Wellcome Open Research, providing they have passed peer review. Preprints, i.e. complete manuscripts that have been submitted to a preprint repository or service (e.g. biorxiv, PeerJ Preprints, arxiv, SocArXiv or PsyArXiv), can be included only if they have a permanent identifier such as a DOI or arxiv identifier. Total number of peer-reviewed publications which you have authored/coauthored. Please exclude abstracts and literature reviews. You should include here systematic reviews and meta analyses. Page 3 of 5

Current and recent research funding (including Wellcome Trust grants) Please list all held in the last five years and any key prior grants (list the most recent first). State the name of the awarding body, name(s) of grantholder(s), title of project, amounts awarded, your role in the project, and start and end dates of support. For all active grants, indicate the number of hours per week that are spent on each project. In addition to research grants, please include details of any recurrent or core funding support that you have held in the last five years, including any Wellcome Trust awards. Please state clearly your role in obtaining the awards, for example, whether you held them in your own right as lead applicant, co-applicant, or as part of a consortium. Please state the value of your own component of the award and the percentage of your time spent on the research. Supporting statement from applicant's current supervisor(s) Please upload a letter of support from your current supervisor(s) If you have already completed your PhD studies and are undertaking postdoctoral research, please provide a letter of support from your present Head of Department. The letter of support should be uploaded to the system and should show clearly the supervisor's (or head's) name, position and address. If there is more than one letter of support, please submit these as a single PDF. Sponsor and research sponsor(s) Details of sponsor at proposed administering organisation The sponsor should be a person you are proposing to work with at the administering organisation. 1 Name, including title (e.g. Professor, Dr) Current position Department Organisation Supporting statement from sponsor Page 4 of 5

Please upload a letter of support from your proposed sponsor. As a guideline, this should be a maximum of 500 words. Have you identified research sponsor(s) that you propose to work with? You will require a research sponsor in every different research environment that you are proposing to work in, but do not have to identify them at this stage. Name and organisation of research sponsor(s) Outline of research programme Please outline your proposed research, including key goals, and indicate the reasons for choice of research location(s). (700 words maximum) You may provide your answer to this question in text entry format or as a PDF attachment. If you are uploading your programme outline, the uploaded document must be in 11 point Arial font and portrait format. Please note, references are not expected at this stage. If you do include references, they will contribute towards the word limit. (700 words max.) Page 5 of 5