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Application summary Type of funding requested Application title Proposed duration of funding (weeks) Proposed start date For Meeting Fellowships, please provide the proposed meeting date. Are you applying through an organisation? Name of administering organisation Lead applicant's address where the grant will be held Department/Division Organisation Street City/Town Postcode/Zipcode Country Details of organisation you plan to visit Lead applicant Lead applicant details Page 1 of 10

Full Name Department Division Organisation Address Line 1 City/Town Postcode Country Telephone No. Email Address ORCID id ORCID id Career history (current/most recent first) From To Position Organisation Education/training From To Qualification Subject Organisation Source(s) of personal salary support Your current work We would like to know about you and the work you are currently doing. What do you do on a daily basis? What are your specific research interests? How will this award fit into your work and career plan? (300 words max.) Clinical status Do you have a medical/veterinary degree? Please specify Page 2 of 10

Are you clinically active? What is your specialty? Please specify Career breaks Have you had any career breaks or periods of part-time work, for example parental or long-term sick leave? Please provide details Peer-reviewed publications and other research outputs List up to ten key prior publications, including original research publications and other scholarly contributions, and other research outputs, e.g. patents. Please include a summary of your contribution to the work associated with each (e.g. developing the idea, conducting the research, supervising staff, writing the paper). For original research publications please provide a DOI, PubMed Central ID or URL. Publications should be in chronological order with the most recent first. Please give citation in full, including title of paper and all authors*. (*All authors, unless more than 10, in which case please use et al, ensuring that your position as author remains clear.) Total number of peer-reviewed publications which you have authored/coauthored. Please exclude abstracts and literature reviews. Are you a named author on any Wellcome Trust funded original peer-reviewed research papers, published from October 2009 onwards? Are all your Wellcome Trust funded original peer-reviewed research papers, published from October 2009 onwards, compliant with our open access policy? Page 3 of 10

Collaborators Will you require any key collaborators for this proposal? Please list up to five key collaborators* (name and organisation) and provide a very brief outline of their role in the proposed activity. *The collaborators named may be replaced with suitable alternatives should it be necessary or appropriate to do so. I confirm that the collaborators named above have agreed to be involved, as described, in the proposed activity and are willing for their details to be included as part of this application. Proposal summary Please provide a summary of your proposal, including key goals. (200 words max.) Details of proposal In this section we would like to know about the purpose of your trip and how it relates to the work of the GFBR. Please describe: (a) the objectives of your trip; (b) why you have chosen the host organisation; (c) what you plan to do during your time (e.g. an itinerary); (d) what outputs you expect to produce (e.g. conference presentations, reports, publications), and planned submission date; (e) how this trip fits in with the goals of the GFBR. 800 words maximum. (800 words max.) Page 4 of 10

In this section we would like to know about the purpose of your project and how it relates to the work of the GFBR. Please describe: (a) the objectives of your project; (b) why you have chosen the topic; (c) what you plan to do during your time on the grant (e.g. meetings, research); (d) what outputs you expect to produce (e.g. conference presentations, reports, publications), and planned submission date; (e) how this project fits in with the goals of the GFBR. 800 words maximum. (800 words max.) In this section we would like to know about the purpose of your meeting and how it relates to the work of the GFBR. Please describe: (a) the objectives of your meeting; (b) who you plan to invite to the meeting; (c) what you plan to do in the meeting (e.g. a draft agenda, key topics of discussion); (d) what outputs you expect to produce (e.g. reports, publications, network development) and, where relevant, planned submission date; (e) how this meeting fits in with the goals of the GFBR. 800 words maximum. (800 words max.) Supporting statement Please provide a signed letter from your supervisor or management at your current organisation, supporting your proposal and confirming that it is necessary and relevant. Page 5 of 10

Supporting statements Please provide a signed letter from your supervisor or management at your current organisation, supporting your trip and confirming that it is necessary and relevant. Please provide a letter from the receiving organisation confirming your invitation and explaining who will be available to support/mentor you during your Fellowship. Data management and sharing Will the proposed research generate data or software outputs that hold significant value as a resource for the wider research community? Please provide a data management and sharing plan (150 words max.) Public engagement Do you have plans for engaging with the non-academic public about your work? Please provide a brief outline of your public engagement plans (250 words max.) Please note that we provide support for Wellcome Trust funded researchers to engage with the non-academic public. Do you wish to receive information about training, funding and other public engagement opportunities? Costs requested and justification Please select the currency in which you wish to apply. Page 6 of 10

Is the selected currency your local currency? What is your local currency? Please state clearly the reasons for requesting costs in the selected currency (100 words max.) Salaries Are you requesting salaries? Salaries / Stipends Description Total Materials and consumables Are you requesting materials and consumables? Materials and consumables Description Total Travel and subsistence Are you requesting travel and subsistence? Travel and subsistence Description Total Miscellaneous costs Are you requesting miscellaneous costs? Miscellaneous other Description Total Page 7 of 10

Justification for resources requested Please provide a brief justification for the resources requested. (300 words max.) Research involving human participants, human biological material and identifiable data Does your project involve human participants, human biological material, or identifiable/potentially identifiable data? Please confirm that you have read the Trust s guidance on the feedback of health-related findings in research and that you are in the process of considering your approach to this. Please state by whom and when the ethics of the project has been, or will be, reviewed and specify any other regulatory approvals that have been obtained, or will be sought. We reserve the right to see relevant approval documents at any point during the lifetime of the grant, in accordance with our policy position on research involving human participants. In the course of your project, do you propose to use facilities within the UK National Health Service (NHS) or to involve patients being cared for by the NHS? Is a formal sponsor required for the project, for example under the Medicines for Human Use (Clinical Trials) Regulations or the Research Governance Framework for Health and Social Care and equivalent guidance? Please indicate which organisation(s) has/have agreed to fulfil this role. Please note that the Wellcome Trust cannot act as sponsor. Page 8 of 10

Freedom to operate/conflicts of interest Describe any freedom to operate issues or potential conflicts of interest that have been identified or that might arise and how these will be or have been addressed. In particular, please consider the following: Do any of the individuals involved in the project hold any consultancies or equities in, or directorships of, companies or other organisations that might have an interest in the results of the proposed activity? Will the proposed activity use technology, materials or other inventions that are subject to any patents or other form of intellectual property protection? Will any element of the activity be subject to agreements with commercial, academic or other organisations, including arrangements with collaborators named in the grant application, that might lead to intellectual property issues or restrictions? (350 words max.) Further information Please use this section to include any further information you would like to share that is relevant to your application. This is optional. Wellcome Trust supported facilities Will the project be based in one of the following Wellcome Trust supported facilities: the Wellcome Trust Sanger Institute a Wellcome Trust Centre Page 9 of 10

a Major Overseas Programme the Francis Crick Institute? Please specify Page 10 of 10